Jirokichi the Rat is wanted in Edo for robbing the mansions of the rich. While on the run, he falls for Osen, a girl who has been sold to a brothel by her criminal brother, Nikichi. When he goes to find Nikichi, Jirokichi immediately falls for Okino, a poor girl who is also under threat from Nikichi.

In Depression-era Tokyo, a struggling middle-aged single father with a young son comes across a homeless young lady and convinces a bar owner to take her in.

In late 19th century Tokyo, Kikunosuke Onoue, the adopted son of a legendary actor, himself an actor specializing in female roles, discovers that he is only praised for his acting due to his status as his father's heir. Devastated by this, he turns to Otoku, a servant of his family, for comfort, and they fall in love. Kikunosuke becomes determined to leave home and develop as an actor on his own merits, and Otoku faithfully follows him.