故事舞台设定在昭和30年代左右的日本，正值高度经济成长期，当时马戏团作为主要娱乐形式，已经融入了人们的日常生活，许多马戏团在日本各地巡回演出，并为了能够参加只有顶级马戏团才被允许参赛的世界级盛会，彼此之间不断竞争、磨炼技艺，在这样的时代背景下，由团长麻利亚率领的向日葵马戏团一边常年为资金短缺而苦恼，一边艰难地继续着巡回演出，就在这时，马戏天才少女鹤卷瑞佳因某些原因来到马戏团，故事也由此开始展开

In a mythical kingdom, the mighty Imperial Knights harness a magical substance known as Aer to power their weapons and protect humanity from the monsters of the forest. But something strange is afoot; the Aer is somehow changing, causing the wilderness to waste away and stirring the woodland beasts to attack with greater frequency. As danger creeps steadily closer to civilization, two young recruits - Flynn, the rigid son of a fallen hero, and the rebellious and brash Yuri - must ride with their fellow Imperial Knights to distant ruins in hopes of uncovering the truth behind the transforming Aer.