Throughout his life Edward Bloom has always been a man of big appetites, enormous passions and tall tales. In his later years, he remains a huge mystery to his son, William. Now, to get to know the real man, Will begins piecing together a true picture of his father from flashbacks of his amazing adventures.

The prehistoric Croods family live in a particularly dangerous moment in time. Patriarch Grug, his mate Ugga, teenage daughter Eep, son Thunk, and feisty Gran gather food by day and huddle together in a cave at night. When a more evolved caveman named Guy arrives on the scene, Grug is distrustful, but it soon becomes apparent that Guy is correct about the impending destruction of their world.

Villainous Gru lives up to his reputation as a despicable, deplorable and downright unlikable guy when he hatches a plan to steal the moon from the sky. But he has a tough time staying on task after three orphans land in his care.