Lightning McQueen, a hotshot rookie race car driven to succeed, discovers that life is about the journey, not the finish line, when he finds himself unexpectedly detoured in the sleepy Route 66 town of Radiator Springs. On route across the country to the big Piston Cup Championship in California to compete against two seasoned pros, McQueen gets to know the town's offbeat characters.

Bob Parr has given up his superhero days to log in time as an insurance adjuster and raise his three children with his formerly heroic wife in suburbia. But when he receives a mysterious assignment, it's time to get back into costume.