故事以主角的第一人称进行，“我”（浅沼晋太郎 配音）是京都大学的一名三年级生，原本期待着有一个玫瑰色多姿多彩的大学生活，希望自己能交上飘逸黑长发的漂亮女朋友。然而事与愿违，主人公经历了一场失败的恋爱，身边的朋友也少得可怜。主人公唯一的朋友小津（吉野裕行 配音），长相像一只妖怪，以捉弄情侣为乐。主人公常常唾弃小津，却离不开他。随后主人公认识了师妹明石（坂本真绫 配音），她是个性格冷淡的黑发少女。 正是因为自己当初选择了网球部，才导致大学两年如此失败，于是“我”假想如果自己入学时选择了另外一个社团，会不会经历另外一种截然不同的大学生活呢？本作从平行时空的角度，围绕着主人公重新从大一新生选择社团开始，选择了各种各样不同社团，发展出不一样的青春故事。

First-year high schooler Midori Asakusa loves anime so much, she insists that "concept is everything" in animation. While she spends her time doodling endless ideas and settings in her sketchbook, she hasn't taken the first step to creating anime, insisting that she can't do it alone. After Asakusa's money-loving best friend Sayaka Kanamori notices her genius and drive – and when it becomes clear that their classmate and charismatic fashion model Tsubame Mizusaki wants to be an animator – the energetic trio start an animation club. Together, the three aim to realize the "ultimate world" that exists in their minds, as they come to see the power that fiction and imagination have on their lives and the world around them.