Universal Century 0079. Humanity has turned even space itself into a battlefield, and the Principality of Zeon forces advance after wiping out half the world's population by carrying out the Operation British (colony drop). In response, the Earth Federation Forces mobilize their overwhelming fighting strength to regain the advantage. The complex intrigues of the Zabi family... Sayla Mass, as she contends with the whims of fate... Hamon and Ramba Ral, who is now a pilot in the Zeon forces... Amuro and Fraw, leading peaceful lives at Side 7... a dark shadow falls across them all. And the Zeon ace Char Aznable, driven by revenge, goes into action as the "Battle of Loum" finally begins.

The episode will focus on the young Zeon ace Char Aznable earning his "Red Comet" nom de guerre, as well as the beginnings of the Federation's "Project V" weapons project that will eventually birth the RX-78-02 Gundam.

《宇宙战舰大和号2199》的续集第四章《宇宙战舰大和号 爱的战士们 第四章 天命篇》确定将于2018年1月27日上映。