真历71年，挺过了多尔西亚攻击的77模组终于抵达月球，而他们也目指近在咫尺的地球，渴望回到吉奥尔人的故国。随着操控Valvrave的次数不断增多，时缟晴人（逢坂良太 配音）饱受符文力量的折磨，最终他和艾尔艾尔弗（木村良平 配音）缔结契约，决心背负着诅咒走到战争的尽头。此后不久，晴人和流木野咲（户松遥 配音）、犬冢久间（小野友树 配音）、山田雷蔵（中村悠一 配音）、连坊小路里见（浪川大辅 配音）、连坊小路晶（悠木碧 配音）驾驶各自的Valvrave成功抵达地球，然而相关数据的偏差却让他们深陷多尔西亚大军的重围。 悲剧的战争与宿命，少男少女充满血和泪的革命之路……

战争终结之后。米凯尔和琪琪一同踏上了新的的旅程。米凯尔此时已经退伍，受08小队其他成员之托，四处打听失踪的队长希罗和艾娜的下落……

It is the year 0083 of the Universal Century. The rebellious Principality of Zeon has been defeated in the One Year War by the Earth Federation. However, a faction of Zeon remnants led by Aguille Delaz fled from the final battle, hiding themselves away. After three long years, they attempt to rise up once more, sending Delaz’s ace pilot, Anavel Gato, to infiltrate a Federation research base to steal one of two secretly developed prototype Gundams along with its deadly nuclear warhead. Threatened by the rogue Gundam suit and seeking to retain peace, the Earth Federation mobilizes the newly developed Albion carrier to recover the stolen unit. Manned by the remaining test pilots, with rookie pilot Kou Uraki piloting the remaining prototype Gundam, the Albion and her crew are determined to stop Gato, retake the stolen Gundam, and prevent the Zeon remnants from starting another war.

U.C. 0083 - Three years after the end of the catastrophic One Year War, peace on Earth and the colonies is shattered by the presence of the Delaz Fleet, a rogue Zeon military group loyal to the ideals of the late dictator Gihren Zabi. Delaz Fleet's ace pilot Anavel Gato, once hailed as "The Nightmare of Solomon", infiltrates the Federation's Torrington base in Australia and steals the nuclear-armed Gundam GP02A "Physalis" prototype. Rookie pilot Kou Uraki - with the aid of Anaheim Electronics engineer Nina Purpleton and the crew of the carrier Albion - pilots the Gundam GP01 "Zephyranthes" prototype in an attempt to recover the stolen Gundam unit and prevent another war from breaking out.

“东京决战”一年后，布里塔尼亚少年鲁路修在11区（原日本国）过着平凡的学生生活。但是，鲁路修与弟弟罗洛的一次出行，遇到了黑色骑士团的余党。在与少女C.C再次结成契约之后，尘封的记忆摆在了鲁路修的面前。 身为帝国王子的鲁路修，为了建立人人平等的世界、让妹妹娜娜丽幸福的世界，而使用GEASS，令人绝对服从的力量，带领黑色骑士团向帝国宣战。带上假面化名ZERO的他，却在一年前的“东京决战”中被好友朱雀击败，被帝国皇帝抹去了记忆。 现在，恢复记忆的鲁路修不仅要面对帝国的强大军事力量，更要面对虚假的弟弟罗洛、失踪的妹妹娜娜丽、不知敌友的中华联盟、内部出现分歧的黑色骑士团。面对内忧外患，鲁路修走上了GEASS之力的诅咒——孤独的王之路。