After a series of mighty clashes with Upper Rank Demons, the Ubuyashiki clan prepares for one last battle with the hellish forces of Muzan Kibutsuji. In order to finally defeat the Demon leader once and for all, the clan devises a training camp for the Demon Slayer Corps, one led by the remaining Hashira - the most elite warriors in the organization. Each Hashira forms a specialized exercise that will hone both their own abilities and the skills of the ordinary soldiers.

Tanjiro Kamado, a boy at the heart of the brewing conflict, recovers from wounds received in a recent fight. While his half-Demon sister Nezuko is studied by researchers like Shinobu Kochou, Tanjiro embarks to train with the Hashira, seeking mastery in each of their assigned areas of expertise to be best prepared for the coming war - skills vital to Tanjiro, as he has vowed to be the very warrior who will eliminate Muzan for good.