In Nazi-occupied France during World War II, a group of Jewish-American soldiers known as "The Basterds" are chosen specifically to spread fear throughout the Third Reich by scalping and brutally killing Nazis. The Basterds, lead by Lt. Aldo Raine soon cross paths with a French-Jewish teenage girl who runs a movie theater in Paris which is targeted by the soldiers.

Through unlocked genetic memories that allow him to relive the adventures of his ancestor in 15th century Spain, Callum Lynch discovers he's a descendant of the secret 'Assassins' society. After gaining incredible knowledge and skills, he is now poised to take on the oppressive Knights Templar in the present day.